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By Paul Haas

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Basel," 1919, 30, 43 ; 1921, 32, 107. See also Ewart: " Jouni. Linn. /' 1896, 31, 364. Wurmser and Jacquot: " Bull. Soc. chim. ," 1923, 5, 305. 3o THE SYNTHESIS OF CARBOHYDRATES The internal temperature of assimilating leaves will vary considerably in accordance with such conditions as the intensity and quality of the light, the character of the leaf surface and so on ; Blackman and Matt had * demonstrated, by thermoelectric means, an excess of 7° C. to 160 C. in the leaves of Primus laurocerasus in bright sunlight above the adjacent shade temperature.

This material is a mixture one component of which is a carbohydrate as is indicated by the facts that it gives the reactions of Molisch and Rubncr, it reduces Benedict's solution and also forms a solid nzazone. The material also contains a more complex substance which reduces Benedict's solution after hydrolysis with acid. In these experiments no free oxygen is given off; it is suggested that it is absorbed on the surface of the powder, possibly forming a peroxide, and this in time may poison the surface and stop the photosynthetic action.

St. "Petersbourg," 191-2t609. f See Palladia: **Ber. deut. bat. Cka," 1891, ip» 229. Mansky: "Itiodteoi. " 1922,132, x8. **Se®Ewart; "Ajia. Bot,/* 1S97, u r 459; 1S9S, 1:2, 379. Pantaaoili: "J&kr. wisa. Bat," 1903, 3^, 167. Urspraiig: *f Ber. d«nt. bot, & " M* 57- INFLUENCE OF ILLUMINATION 31 be lost by reflection, absorption or transmission. Of the energy absorbed by the leaf, many have shown that a small proportion only, and this in varying quantity, is used in carbon assimilation. Owing to ignorance of certain factors and the degree of their significance in the sequence of carbon assimilation, it is not possible to give a satisfactory account of the energy relationships of the plant.

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