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The punctuated equilibrium model (Eldredge and Gould 1972; Gould and Eldredge 1977; Stanley 1979) has had a major impact on the study of Paleozoic faunas, and many studies have confirmed punctuated patterns of change in Paleozoic species. Lieberman et al. (1995) found a pattern of punctuated stasis in brachiopods from the Middle Devonian Hamilton Group of the Appalachian Basin. In this study, we examine contemporary brachiopods from the Traverse Group of the Michigan Basin to see whether the patterns found in the Appalachian Basin are regional or local in scope.

4) could have been in response to their adapting to changing paleoenvironmental conditions prevailing in those time periods. 028), which is representative of little or no morphological change, as expected in an EEU. 05), concurs with the current classification system in atrypides. 4). This suggests that this difference may be either due to differences in sample size (as the between-genera distances are based on several samples of the same family, whereas the distances through time are based on sub-samples of the same genus), or it may be due to numerous real short-term changes within the lineages, which get averaged out when comparisons are made between genera.

0 Procrustes units. 8 Procrustes units. 3 Procrustes units. Overall, geographic variation within genera is greater than temporal variation. 028 Procrustes units respectively). 037 Procrustes units for Middle Devonian respectively). 040). 5). 01). 1 Taxonomy In this study, various atrypid genera from the Silurian-Devonian atrypid subfamilies were identified based on phenotypic characters other than shell shape. Overall, significant morphometric differences exist in the shells between subfamilies, thus supporting suggestions put forward by Copper (1973) that these groups are distinct.

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